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Apocleinae
Amblyonychus
Eccritosia
Furcilla
Mallophora
Megaphorus
Nevadasilus
Proctacanthella
Proctacanthus
Promachella
Promachus
Efferia-group
Albibarbefferia
Aridefferia
Carinefferia
Efferia *
Nerax
Pogonioefferia
Triorla
Tuberculefferia

Efferia Coquillet, 1893
[Apocleinae: Efferia-group]

Neotropical Nearctic Palaearctic Afrotropical Oriental Australian & Oceania
  typ.loc.        

Description: Coquillett, D.W. (1893): A new asilid genus related to Erax. - The Canadian Entomologist 25: 175-177 [175]; Ottawa.
Type species: Efferia candida Coquillett, 1893: 176
Literature: Coquillett, D.W. (1893): A new asilid genus related to Erax. - The Canadian Entomologist 25: 175-177; Ottawa [key]. Hine, J.S. (1919): Robber flies of the genus Erax. - Annals of the Entomological Society of America 12(2): 103-157, pls. 7-9; Columbus [key]. Wilcox, J. (1966): Efferia Coquillett in America North of Mexico (Diptera, Asilidae). - Proceedings of the California Academy of Sciences 34: 85-234; San Francisco [key]. Parks, L. (1968): Synopsis of robberfly genera allied to Efferia and Eicherax, including a new genus. - Pan-Pacific Entomologist 44: 171-179; San Francisco [key]. Forbes, G.S. (1988): Three new species of Efferia from southern California and Arizona (Diptera: Asilidae). - Annals of the Entomological Society of America 81: 554-559; Columbus.

Nearctic species:

  1. Efferia anacapai (Wilcox & Martin, 1945)
  2. Efferia anomala (Bellardi, 1861)
  3. Efferia antiochi Wilcox, 1966
  4. Efferia azteci Wilcox, 1966
  5. Efferia bryanti Wilcox, 1966
  6. Efferia candida Coquillett, 1893
  7. Efferia completa (Macquart, 1838)
  8. Efferia davisi Wilcox, 1966
  9. Efferia gila Wilcox, 1966
  10. Efferia halli Wilcox, 1966
  11. Efferia imperialis Forbes, 1988
  12. Efferia luna Wilcox, 1966
  13. Efferia macroxipha Forbes, 1988
  14. Efferia pernicis Coquillett, 1893
  15. Efferia tricella (Bromley, 1951)

Efferia candida Coquillett, 1893

- male -

dorsal view   lateral view
head dorsal   head lateral
head frontal   ocellar tubercle
scutum dorsal   scutum lateral
anterior scutum lateral   scutum forntal
scutellum dorsal   scutellum lateral
wing   wing
wing   abdomen dorsal
abdomen lateral   hypopygium dorsal
hypopygium lateral

- female -

dorsal view   lateral view
wing   ovipositor lateral


Key to the species

Founded on: Wilcox, J. (1966): Efferia Coquillett in America North of Mexico (Diptera, Asilidae). - Proceedings of the California Academy of Sciences 34: 85-234; San Francisco. [amended]

1 Males. 2
- Females. 16

2 Abdomen with long white parted hair directed outward on 3 or more segments. 3
- Abdomen without long paired hair. 12

3 Femora reddish posteriorly. Usually larger flies, 25-32 mm. 4
- Femora black, at most narrowly reddish at the base or apex. Smaller flies, 13-25 mm. 6

4 Tergites 2-3 dark brown basally, 2-5 with long white parted hair becomeing shorter on apical segments. Bristles of palpi and occipput, and hairs on hypopygium largely black. 6-10 black scutellar bristles. Length 25-30 mm . . . . . Efferia anomala (Bellardi, 1861) [MEX, KS?, TX, AZ, CA]  
- Abdomen wholly silvery white pollinose, segments 2-5 with white parted hair which becomes shorter with each segment apically. Bristles of palpi and occiput, and hairs of hypopygium usually, largely white. 5

5 Scutellar bristles black. Thorax distinctly reddish. Length 25-32 mm . . . . . Efferia tricella (Bromley, 1951) [AZ, NM, MEX, SON]  
- Scutellar bristles whitish. Thorax reddish black. Length 19-26 mm . . . . . Efferia macroxipha Forbes, 1988 [CA]  

6 4th posterior branch of radius (R4) ends in the 2nd and 3rd posterior branch of radius (R2+3). Hypopygium usually yellowish red. Tergites 2-7 with white parted hair. Length 13-19 mm . . . . . Efferia azteci Wilcox, 1966 [AZ, CA]  
- 4th posterior branch of radius (R4) ends in the costa. Hypopygium usually black. 7

7 Scutellum with dense clumps of white hairs on either side of the disc, about 16 black marginal bristles. Mystax with many black bristles, white hairs intermixed. tergites 2-7 with white parted hairs decreasing in length on 4-7. Length 21-24 mm . . . . . Efferia anacapai (Wilcox & Martin, 1945) [CA]  
- Scutellum with scattered hairs on the disc, usually not more than 8 marginal bristles. Mystax wholly black. 8

8 Hypopygium dorsally black haired and usually some black hairs on lower margin of epandria and in the fringe on gonocoxite. Scutellar and postocular bristles all or in part black. 9
- Hypopygium white haired, sometimes a few black hairs basally at middle of epandria. Scutellar and postocular bristles white. 11

9 Parted white hairs on abdomen becoming progressively shorter from segment 2 to 5, short appressed white on 6-7. Tail of gonocoxite slender and as long as basal section. Length 21-25 mm . . . . . Efferia luna Wilcox, 1966 [NM, KS, TX]  
- Parted white hairs on tergites 2-5 subequal in length, long appressed on 6-7. 10

10 Hairs on the frons all or in part black. Tail of gonocoxite about half length of basal section. hairs sternites 1-7 erect but becoming shorter apically. Length 15-22 mm . . . . . Efferia halli Wilcox, 1966 [CA]  
- Hairs on the frons white. Tail of gonocoxite about two-thirds length of basal section. Sternites 1-5 with erect hairs, short on 6-7. Length 21-24 mm . . . . . Efferia gila Wilcox, 1966 [AZ]  

11 Hairs on sternites short, subequal in length to pedicel. Upper hairs in mystax subequal in length to scape and pedicel combined. length 16-21 mm . . . . . Efferia davisi Wilcox, 1966 [AZ, NV, NM, UT, MEX]  
- Hairs on sternites 1-4 erect and at least as long as scape and pedicel combined. Upper hairs in mystax as long as scape, pedicel, and postpedicel combined. Length 16-25 mm . . . . . Efferia candida Coquillett, 1893 [CA, AZ, NM, KS?, OK?, MEX]  

12 Tergites 2-6 broadly black basally and grayish white pollinose on apical one-fourth to one-fifth, giving an annulated appearance to abdomen, tergite 7 entirely white pollinose, sides of tergites and sternites grayish white pollinose. Lower lobe of gonocoxite short . . . . . . Efferia imperialis Forbes, 1988 [CA, AZ, MEX, SON]  
- Other combination of characters. 13

13 Abdomen uniformly gray pollinose. Femora usually reddish at base ans apex. 4th posterior branch of radius (R4) usually ends in the 2nd and 3rd posterior branch of radius (R2+3). 8-10 scutellar brsitles, usually white. Length 15-24 mm . . . . . Efferia pernicis Coquillett, 1893 [CA, AZ, UT, NV]  
- Tergites 2-5 largely black, 6-7 densely white pollinose. 14

14 Abdomen usually wholly black haired, sometimes with a few white hairs laterally on segments 1-3. Hypopygium black haired. Femora black. Length 16-20 mm . . . . . Efferia completa (Macquart, 1838) [TX, MEX]  
- Abdomen with numerous white hairs laterally at least on segments 1-3. Hypopygium largely white haired. 15

15 White hairs on abdomen confined to segments 1-3. Femora usually reddish brown dorsally. Length 17-24 mm . . . . . Efferia bryanti Wilcox, 1966 [AZ]  
- Abdominal segments 1-4 with white hairs. Femora wholly black. Length 20-26 mm . . . . . Efferia antiochi Wilcox, 1966 [CA]  

16 Ovipositor 6.6. to 7.2 mm long. 17
- Ovipositor at most 5.4 mm long. 19

17 Scutellum with sparse erect black and white hairs. Ovipositor 6.6 mm long . . . . . Efferia antiochi Wilcox, 1966 [CA]  
- Scutellum with dense clumps of white hairs on the disc. Ovipositor longer. 18

18 Scutellar bristles black. Ovipositor 7.2 mm long. . . . . Efferia anacapai (Wilcox & Martin, 1945) [CA]  
- Scutellar bristles whitish. Ovipositor 8.1-9.4 mm long . . . . . Efferia macroxipha Forbes, 1988 [CA]  

19 4th posterior branch of radius (R4) usually ends in or nearby the 2nd and 3rd posterior branch of radius (R2+3). 20
- 4th posterior branch of radius (R4) usually ends in the costa. 22

20 Base and apex of the fomora reddish. Ovipositor 4.5-5.0 mm long . . . . . Efferia pernicis Coquillett, 1893 [CA, AZ, UT, NV]  
- Femora wholly black. 21

21 Cerci of ovipositor without distinct spines. Ovipositor 2.5-3.0 mm long . . . . . Efferia azteci Wilcox, 1966 [AZ, CA]  
- Cerci of ovipositor spinose. Ovipositor 3.7-4.3 mm long. . . . . . . Efferia imperialis Forbes, 1988 [CA, AZ, MEX, SON]  

22 Posterior side or dorsum of the femora reddish. 23
- Femora wholly black. 25

23 Dorsum of femora reddish. Ovipositor 3.0-3.5 mm long, with short spines on the tip . . . . . Efferia bryanti Wilcox, 1966 [AZ]  
- Posterior side of femora reddish. Ovipositor with hairs at tip. 24

24 Abdomen wholly whitish pollinose. 4 scutellar brsitles, white or black. Palpi white haired. Ovipositor 4.5-5.0 mm long . . . . . Efferia tricella (Bromley, 1951) [AZ, NM, MEX, SON]  
- Abdominal segments blackish basally. 8 black scutellar bristles. Palpi black haired. Ovipositor 4.5-5.0 mm long . . . . . Efferia anomala (Bellardi, 1861) [MEX, KS?, TX, AZ, CA]  

25 Ovipositor forked at tip. Hairs on anterior scutum as long as scape and pedicel combined. Ovipositor 3.5-4.0 mm long . . . . . Efferia completa (Macquart, 1838) [TX, MEX]  
- Ovipositor not divided at tip. Hairs on anterior scutum not longer than pedicel. 26

26 Scutellar and postocular bristles white. Abdomen densely white pollinose and white haired, segment 7 sometimes thinly pollinose and black haired dorsally. 27
- Scutellar and postocular bristles all or in part black. Abdomen white pollinose, segments 6-7 thinly pollinose and with black hairs at least dorsally. 28

27 Sternites with all short white hairs. Hairs above in mystax quite dense but not longer than scape. Ovipositor 3.7 mm long . . . . . Efferia davisi Wilcox, 1966 [AZ, NV, NM, UT, MEX]  
- Sternites 1-3 with long erect hairs. Hairs above in mystax dense and as long as or longer than scape and pedicel combined. Ovipisitor 3.6-3.7 mm long . . . . . Efferia candida Coquillett, 1893 [CA, AZ, NM, KS?, OK?, MEX]  

28 Discal hairs of scutellum as long as scape and pedicel combined. Lateral hairs on the frons in part black. Face 0.9 width of one eye. Ovipositor 3.3. mm long . . . . . Efferia halli Wilcox, 1966 [CA]  
- Discal hairs of scutellum not longer than pedicel. Lateral hairs on the frons white. 29

29 Fore and middle tibiae and tarsomeres reddish. Face at antennae 0.8 width of one eye. Ovipositor 4.2 mm long . . . . . Efferia luna Wilcox, 1966 [NM, KS, TX]  
- Fore and middle tibiae at the tips and the tarsomeres black. Face at antennae 0.9 width of one eye. Ovipositor 5.4 mm long . . . . . Efferia gila Wilcox, 1966 [AZ]  

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

Contents: © Fritz Geller-Grimm, Jorge N. Artigas, 2007
Layout & images: © Fritz Geller-Grimm, 2007